Cimolodonta

Cimolodonta Temporal range: Cretaceous-Eocene
A reconstruction of Ptilodus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Order:	Multituberculata
Suborder:	Cimolodonta
Superfamilies
Paracimexomys group Djadochtatherioidea Ptilodontoidea Taeniolabidoidea Other families Boffiidae Cimolomyidae Eucosmodontidae Kogaionidae Microcosmodontidae

The Cimolodonta are a taxon of extinct mammals that lived from the Cretaceous to the Eocene. They were some of the more derived members of the extinct order Multituberculata. They probably lived something of a rodent-like existence until their ecological niche was assumed by true rodents. The more basal multituberculates are found in a different suborder, "Plagiaulacida".

Cimolodonta is apparently a natural (monophyletic) suborder. Remains have been identified from across the Northern Hemisphere. The taxon is recognized as the informal Paracimexomys group and the superfamilies Djadochtatherioidea, Taeniolabidoidea, and Ptilodontoidea. Additionally, and of uncertain affinities, are the families Cimolomyidae, Boffiidae, Eucosmodontidae, Kogaionidae, Microcosmodontidae and the two genera Uzbekbaatar and Viridomys. More precise placement of these types awaits further discoveries and analysis.

Taxonomy

Suborder †Cimolodonta McKenna, 1975

• Superfamily Incertae sedis
  • Family Incertae sedis
    • Subfamily Incertae sedis
      • Genus? †Ameribaatar Eaton & Cifelli, 2001
        • Species? †A. zofiae Eaton & Cifelli, 2001
      • Genus? †Bryceomys Eaton, 1995
        • Species †B. fumosus Eaton, 1995
        • Species †B. hadrosus Eaton, 1995
        • Species †B. intermedius Eaton & Cifelli, 2001
      • Genus †Cedaromys Eaton & Cifelli, 2001
        • Species †C. bestia (Eaton & Nelson, 1991) Eaton & Cifelli, 2001 [=Paracimexomys? bestia Eaton & Nelson, 1991]
        • Species †C. parvus Eaton & Cifelli, 2001
      • Genus? †Dakotamys Eaton, 1995; E. Cret. CNA.
        • Species? †D. sp. (Utah, USA) Eaton, 1995
        • Species †D. malcolmi Eaton, 1995
      • Genus? †Fractimus Higgins, 2003
        • Species? †F. palmorum Higgins, 2003
      • Genus †Halodon Marsh, 1889
        • Species †H. serratus Marsh, 1889
      • Genus †Ptilodus (Marsh, 1889) Gidley, 1909
        • Species †P. serratus (Marsh, 1889) Gidley, 1909
      • Genus? †Uzbekbaatar Kielan-Jaworowska & Nesov, 1992
        • Species? †U. kizylkumensis Kielan-Jaworowska & Nesov, 1992
      • Genus? †Barbatodon Rãdulescu & Samson, 1986
        • Species? †B. transylvanicum Rãdulescu & Samson, 1986
• Paracimexomys group Archibald, 1982
  • • • Genus †Paracimexomys Archibald, 1982
        • Species? †P. crossi Cifelli, 1997
        • Species? †P. magnus (Sahni, 1972) Archibald, 1982 [Cimexomys magnus Sahni, 1972]
        • Species? †P. magister (Fox, 1971) Archibald, 1982 [Cimexomys magister Fox, 1971]
        • Species? †P. perplexus Eaton & Cifelli, 2001
        • Species? †P. robisoni Eaton & Nelson, 1991
        • Species? †P. priscus (Lillegraven, 1969) Archibald, 1982 [Cimexomys priscus Lillegraven, 1969; genotipo Paracimexomys sensu Eaton & Cifelli, 2001]
        • Species †P. propriscus (Lillegraven, 1969) Hunter et Al., 2010
      • Genus? Cimexomys Fox, 1971
        • Species? †C. antiquus Fox, 1971
        • Species? †C. gregoryi Eaton, 1993
        • Species? †C. electus
        • Species? †C. bellus
        • Species? †C. clarki
        • Species? †C. judithae Sahni, 1972 [Paracimexomys? judithae (Sahni, 1972) Archibald, 1982]
      • Genus Cimexomys Sloan & Van Valen, 1965 sensu stricto?
        • Species? †C. arapahoensis
        • Species †C. minor Sloan & Van Valen, 1965
        • Species? †C. gratus (Jepson, 1930) Lofgren, 1995 [Cimexomys hausoi Archibald, 1983; Eucosmodon gratus Jepson, 1930; Mesodma? ambigua? Jepson, 1940; Stygimus gratus (Jepson, 1930)]
  • Family †BoffidaeHahn & Hahn, 1983
    • • Genus †Boffius Vianey-Liaud, 1979
        • Species †Boffius splendidus Vianey-Liaud, 1979 [Boffiidae Hahn & Hahn, 1983 sensu Kielan-Jaworowska & Hurum, 2001]
  • Family Cimolomyidae Marsh, 1889 sensu Kielan-Jaworowska & Hurum, 2001
    • • Genus Essonodon Simpson, 1927
        • Species? †E. browni Simpson, 1927 [cimolodontidae? Kielan-Jaworowska & Hurum (2001)]
      • Genus Buginbaatar Kielan-Jaworowska & Sochava, 1969
        • Species? †B. transaltaiensis Kielan-Jaworowska & Sochava, 1969
      • Genus Meniscoessus Cope, 1882 [Dipriodon Marsh, 1889, Tripriodon Marsh, 1889, Selenacodon Marsh, 1889, Halodon Marsh, 1889, Oracodon Marsh, 1889]
        • Species †M. caperatus Marsh, 1889
        • Species †M. collomensis Lillegraven, 1987
        • Species †M. ferox Fox, 1971a
        • Species †M. intermedius Fox, 1976b
        • Species †M. major (Russell, 1936)
        • Species †M. greeni
        • Species †M. conquistus Cope, 1882
        • Species †M. robustus (Marsh, 1889)
        • Species †M. seminoensis Eberle & Lillegraven, 1998a
        • Species †M. borealis [M. robustus]
      • Genus †Cimolomys Marsh, 1889 [=? Allacodon Marsh, 1889; Meniscoessus; Ptilodus; Selenacodon Marsh, 1889]
        • Species? †C. lentus [Allacodon lentus]
        • Species? †C. pumilus [Allacodon pumilus]
        • Species †C. clarki Sahni, 1972
        • Species †C. gracilis Marsh, 1889 [Cimolomys digona Marsh, 1889; Meniscoessus brevis; Ptilodus gracilis Osborn, 1893; Selenacodon brevis Marsh, 1889]
        • Species †C. trochuus Lillegraven, 1969
        • Species †C. milliensis Eaton, 1993a
        • Species †C. sp.2
• Superfamily Ptilodontoidea Cope, 1887 sensu McKenna & Bell, 1997 e Kielan-Jaworowska & Hurum, 2001
  • Family Cimolodontidae Marsh, 1889 sensu Kielan-Jaworowska & Hurum, 2001
    • • Genus †Liotomus Lemoine, 1882
        • Species? †L. marshi (Lemoine, 1882) Cope, 1884 [Neoctenacodon marshi Lemoine, 1882?] [eucosmodontidae? McKenna & Bell, 1997]
      • Genus †EssonodonSimpson, 1927
        • Species? †E. browni Simpson, 1927 [cimolomyidae? Kielan-Jaworowska & Hurum (2001)]
      • Genus †Anconodon Jepsen, 1940
        • Species? †A. lewisi (Simpson, 1935)
        • Species †A. rarus
        • Species †A. russelli Sloan, 1987
        • Species †A. gibleyi (Simpson, 1935)
        • Species †A. cochranensis (Russell, 1929) [Liotomus russelli (Simpson, 1935); Ectopodon cochranensis Russell, 1967]
      • Genus †Cimolodon Marsh, 1889 [Nanomys Marsh, 1889, Nonomyops Marsh, 1892]
        • Species †C. electus
        • Species †C. nitidus Marsh, 1889 [Allacodon rarus Marsh, 1892 sensu Clemens, 1964a; Nanomys minutus Marsh, 1889; Nonomyops minutus (Marsh, 1889) Marsh, 1892]
        • Species †C. parvus
        • Species †C. similis Fox, 1971
        • Species †C. gracilis Marsh, 1889
        • Species †C. sp.2
      • Genus Neoliotomus Jepsen, 1930
        • Species †N. conventus Jepsen, 1930
        • Species †N. ultimus (Granger & Simpson, 1928)
  • Family Ptilodontidae Cope, 1887 sensu McKenna & Bell, 1997
    • Subfamily Neoplagiaulacidae Ameghino, 1890 [Ptilodontidae: Neoplagiaulacinae Ameghino, 1890 sensu McKenna & Bell, 1997]
      • Genus MesodmaMarsh, 1889
        • Species? †Mesodma sp.1, cf. M. formosa
        • Species? †Mesodma sp.2, cf. M. formosa
        • Species? †Mesodma sp.3, cf. M. formosa
        • Species? †Mesodma sp.4
        • Species? †Mesodma sp.5
        • Species? †M. hensleighi
        • Species? †M. senecta
        • Species? †M. garfieldensis Archibald, 1982
        • Species? †M. pygmaea Sloan, 1987
        • Species †M. formosa (Marsh, 1889) [Halodon formosus Marsh, 1889??]
        • Species †M. primaeva [Perectypodus primaeva]
        • Species †M. thompsoni Clemens, 1964
        • Species †M. ambigua Jepsen, 1940 [†Mesodma. sp.4, cf. Mesodma thompsoni??]
        • Species †M. siberlingi (Simpson, 1935) [Ectypodus?; Mimetodon?]
      • Genus Ectypodus Matthew & Cranger, 1921 [Charlesmooria Kühne, 1969 ]
        • Species? †E. childei (Kühne, 1969)
        • Species? †E. simpsoni (Jepsen, 1930)
        • Species? †E. elaphus Scott, 2005
        • Species †E. aphronorus Sloan, 1981
        • Species †E. szalayi Sloan, 1981
        • Species †E. musculus Matthew & Granger, 1921
        • Species †E. powelli Jepsen, 1940
        • Species †E. tardus (Jepsen, 1930)
      • Genere Mimetodon Jepsen, 1940
        • Species †M. krausei Slaon, 1981
        • Species †M.? nanophus (Holtzman, 1978) [Neoplagiaulax nanophus Holtzman, 1978]
        • Species †M. trovessortianus
        • Species †M. churchilli Jepsen, 1940
        • Species †M. douglassi
      • Genus Neoplagiaulax Lemoine, 1882
        • Species †N. nanophus Holtzman, 1978
        • Species †N. nelsoni Sloan, 1987
        • Species †N. mckennai Sloan, 1987 [N. mckennaiai Sloan, 1987 (lapsus calami)]
        • Species †N. jepi Sloan, 1987
        • Species †N. annae Vianey-Liaud, 1986
        • Species †N. fractus Dorr, 1952
        • Species †N. nicolai Vianey-Liaud, 1986
        • Species †N. macintyrei Slaon, 1981
        • Species †N. hazeni (Jepsen, 1940)
        • Species †N. sp. cf. N. hazeni Krause, 1977, e Fox, 1990
        • Species †N. paskapooensis Scott, 2005 [Neoplagiaulax hunteri Krishtalka, 1973 (in partim); Neoplagiaulax sp. 2 Fox, 1990]
        • Species †N. cimolodontoides Scott, 2005 [Neoplagiaulax sp. 3 Fox, 1990]
        • Species †N. macrotomeus (Wilson, 1956)
        • Species †N. copei Lemoine, 1885
        • Species †N. eocaenus (Lemoine, 1880) [genotipo]
        • Species †N. grangeri (Simpson, 1935)
        • Species? †N. serrator Scott, 2005 [N. hunteri Krishtalka, 1973 (in partim); N. sp. 1 Fox, 1990]
        • Species †N. kremnus Johnston & Fox, 1984
        • Species †N. sp. cf. N. hunteri (Simpson, 1936)
        • Species †N. hunteri (Simpson, 1936)
      • Genus †Parectypodus Jepsen, 1930
        • Species? †P. lovei Sloan, 1966
        • Species? †P. vanvaleni Sloan, 1981
        • Species? †P. corystes Scott, 2003
        • Species †P. sloani Schiebout, 1974
        • Species †P. trovessartianus (Cope, 1882) [P. trouessarti; Ptilodus; Mimetodon; Neoplagiaulax]
        • Species †P. sylviae (Rigsby, 1980) [Ectypodus sylviae Rigby, 1980]
        • Species †P. pattersoni Sloan, 1987
        • Species †P. armstrongi Johnston & Fox, 1984
        • Species †P. sinclairi (Simpson, 1935)
        • Species †P. laytoni Jepsen, 1940
        • Species †Parectypodus simpsoniP. simpsoni
      • Genus †Carnaysia Vianey-Liaud, 1986
        • Species †C. manueli Vianey-Liaud, 1986
        • Species †C. davidi Vianey-Liaud, 1986
      • Genus †Krauseia Vianey-Liaud, 1986
        • Species †K. clemensi (Sloan, 1981) [Parectypodus clemensi Sloan, 1981]
      • Genus †XironomysRigby, 1980
        • Species †X. swainae Rigby, 1980 [Xyronomys; ?Eucosmodontidae]
      • Genus †Xancolomys Rigby, 1980
        • Species †X. mcgrewiRigby, 1980
      • Genus †MesodmopsTong & Wang, 1994
        • Species †M. dawsonae Tong & Wang, 1994
    • Subfamily Ptilodontidae Cope, 1887 [Ptilodontidae: Ptilodontinae Cope, 1887 sensu McKenna & Bell, 1997]
      • Genus †Kimbetohia Simpson, 1936
        • Species †K. cambi [Granger, Gregory & Colbert in Matthew, 1937, or Simpson, 1936]
        • Species †K. sp. cf. K. cambi
      • Genus †Ptilodus Cope, 1881 [Chirox Cope, 1884]
        • Species? †P. fractus
        • Species †P. tsosiensis Sloan, 1981
        • Species †P. kummae Krause, 1977
        • Species †P. ferronensis Gazin, 1941
        • Species †P. gnomus Scott, Fox & Youzwyshyn, 2002 [cf. Ectypodus hazeni (Jepsen, 1940) Gazin, 1956]
        • Species †P. mediaevus Cope, 1881 [Ptilodus plicatus (Cope, 1884); Chirox plicatus Cope, 1884]
        • Species †P. nellieae
        • Species †P. sinclairi
        • Species †P. montanus Douglass, 1908 [P. gracilis Gidley, 1909; P. admiralis Hay, 1930]
        • Species †P. wyomingensis Jepsen, 1940
      • Genus †Baiotomeus Krause, 1987
        • Species †B. russelli Scott, Fox & Youzwyshyn, 2002
        • Species †B. douglassi (Simpson, 1935) [Ptilodus; Mimetodon; Neoplagiaulax]
        • Species †B. lamberti Krause, 1987
        • Species †B. rhothonion Scott, 2003
      • Genus †Prochetodon Jepsen, 1940
        • Species? †P. speirsae Scott, 2004
        • Species †P. cavus Jespen, 1940
        • Species †P. foxi Krause, 1987
        • Species †P. taxus Krause, 1987
        • Species? †P. sp. cf. P. cavus Jespen, 1940
      • Genere †Kogaionidae Rãdulescu & Samson, 1996
        • Species †K. ungureanui Rãdulescu & Samson, 1996
      • Genus †Hainina Vianey-Liaud, 1979
        • Species †H. belgica Vianey-Liaud, 1979
        • Species †H. godfriauxi Vianey-Liaud, 1979
        • Species †H. pyrenaica Peláez-Campomanes, López-Martínez, Álvarez-Sierra & Daams, 2000
        • Species †H. vianeyae Peláez-Campomanes, López-Martínez, Álvarez-Sierra & Daams, 2000
    • Subfamily Eucosmodontidae Jepsen, 1940 sensu Kielan-Jaworowska & Hurum, 2001 [Eucosmodontidae: Eucosmodontinae Jepsen, 1940 sensu McKenna & Bell, 1997]
      • Genus †ClemensodonKrause, 1992
        • Species †C. megaloba Krause, 1992 [Kimbetohia cambi, in partim]
      • Genus †Eucosmodon Matthew & Granger, 1921
        • Species? †E. gratus Jepsen, 1940 [= Stygimys gratus??]
        • Species †E. primus [Granger & Simpson, 1929 or Sloan, 1981]
        • Species †E. americanus (Cope, 1885)
        • Species †E. molestus (Cope, 1869) [1886?] [Neoplagiaulax molestus Cope, 1869]
      • Genus †Stygimys Sloan & Van Valen, 1965
        • Species †S. camptorhiza Johnston & Fox, 1984
        • Species †S. gratus (Jepsen, 1930)
        • Species? †S. sp. cf. S. kuszmauli
        • Species †S. kuszmauli [Eucosmodon kuszmauli]
        • Species †S. jepseni (Simpson, 1935) [S. sp. cf. S. kuszmauli]
        • Species †S. teilhardi (Granger & Simpson, 1929)
    • Subfamily Microcosmodontidae Holtzman & Wolberg, 1977 [Eucosmodontidae: Microcosmodontinae Holtzman & Wolberg, 1977 sensu McKenna & Bell, 1997]
      • Genus †PentacosmodonJepsen, 1940
        • Species †P. pronus Jepsen, 1940 [Djadochtatheroid? (Kielan-Jaworowska & Hurum, 2001)]
      • Genus †Acheronodon Archibald, 1982
        • Species †A. garbani Archibald, 1982
      • Genus †Microcosmodon Jepsen, 1930
        • Species †M. conus Jepsen, 1930
        • Species †M. rosei Krause, 1980
        • Species †M. arcuatus Johnston & Fox, 1984
        • Species †M. woodi Holtzman & Wolberg, 1977 [Eucosmodontine?]
        • Species †M. harleyi Weil, 1998
• Superfamily Djadochtatheroidea Kielan-Jaworowska & Hurum, 1997 sensu Kielan-Jaworowska & Hurum, 2001[Djadochtatheria Kielan-Jaworowska & Hurum, 1997]
  • • • Genus? †BulganbaatarKielan-Jaworowska, 1974
        • Species? †B. nemegtbaataroides Kielan-Jaworowska, 1974
      • Genus? †Chulsanbaatar Kielan-Jaworowska, 1974
        • Species? †C. vulgaris Kielan-Jaworowska, 1974 [Chulsanbaataridae]] Kielan-Jaworowska, 1974
      • Genus †Nemegtbaatar Kielan-Jaworowska, 1974
        • Species? †N. gobiensis Kielan-Jaworowska, 1974
  • Family Sloanbaataridae Kielan-Jaworowska, 1974
    • • Genus †Kamptobaatar Kielan-Jaworowska, 1970
        • Species? †K. kuczynskii Kielan-Jaworowska, 1970
      • Genus †Nessovbaatar Kielan-Jaworowska & Hurum, 1997
        • Species †N. multicostatus Kielan-Jaworowska & Hurum, 1997
      • Genus †Sloanbaatar Kielan-Jaworowska, 1974
        • Species †S. mirabilis Kielan-Jaworowska, 1974 [Sloanbaatarinae]
  • Family Djadochtatheriidae Kielan-Jaworowska $ Hurum, 1997
    • • Genus †DjadochtatheriumSimpson, 1925
        • Species †D. matthewi Simpson, 1925
      • Genus †Catopsbaatar Kielan-Jaworowska, 1974
        • Species †C. catopsaloides (Kielan-Jaworowska, 1974) Kielan-Jaworowska, 1994 [Djadochtatherium catopsaloides Kielan-Jaworowska, 1974 ; Catopsalis catopsaloides (Kielan-Jaworowska, 1974) Kielan-Jaworowska & Sloan, 1979]
      • Genus †Tombaatar Kielan-Jaworowska, 1974
        • Species †T. sabuli Rougier, Novacek & Dashzeveg, 1997
      • Genus †Kryptobaatar Kielan-Jaworowska, 1970 [Gobibaatar Kielan-Jaworowska, 1970 , Tugrigbaatar Kielan-Jaworowska & Dashzeveg, 1978]
        • Species †K. saichanensis Kielan-Jaworowska & Dashzeveg, 1978 [Tugrigbaatar saichaenensis Kielan-Jaworowska & Dashzeveg, 1978??]
        • Species †K. dashzevegi Kielan-Jaworowska, 1970
        • Species †K. mandahuensis Smith, Guo & Sun, 2001
        • Species †K. gobiensis Kielan-Jaworowska, 1970 [Gobibaatar parvus Kielan-Jaworowska, 1970 ]
• Superfamily Taeniolabidoidea Granger & Simpson, 1929 sensu Kielan-Jaworowska & Hurum, 2001
  • Family Taeniolabididae Granger & Simpson, 1929
    • • Genus †Catopsalis Cope, 1882
        • Species †C. alexandri Middleton, 1982
        • Species †C. collariensis
        • Species †C. joyneri Sloan & Van Valen, 1965
        • Species †C. matthewi
        • Species †C. waddleae Buckley, 1995
        • Species †C. calgariensis Russell, 1926
        • Species †C. fissidens (Cope, 1884)
        • Species †C. foliatus Cope, 1882
        • Species †C. utahensis Gazin, 1939
      • Genus †Prionessus Matthew & Granger, 1925
        • Species †P. lucifer Matthew & Granger, 1925
      • Genus †Lambdopsalis Chow & Qi, 1978
        • Species †L. bulla Chow & Qi, 1978
      • Genus †Sphenopsalis Matthew, Granger & Simpson, 1928
        • Species †S. nobilis Matthew, Granger & Simpson, 1928
      • Genus †Taeniolabis Cope, 1882
        • Species †T. lamberti Simmons, 1987
        • Species †T. taoensis (Cope, 1882)

References

• Kielan-Jaworowska Z. and Hurum J.H. (2001), "Phylogeny and Systematics of multituberculate mammals". Paleontology 44, p.389-429.
• Much of this information has been derived from [1] Multituberculata Cope, 1884.