Mythicomyiidae

Mythicomyiidae
Undescribed psiloderoidine genus from Australia.
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Suborder:	Brachycera
Infraorder:	Asilomorpha
Superfamily:	Asiloidea
Family:	Mythicomyiidae Melander , 1902
Subfamilies
Psiloderoidinae Platypyginae Glabellulinae Empidideicinae Leylaiyinae Mythicomyiinae

Mythicomyiids are very tiny flies (0.5–5.0 mm) found throughout most parts of the world, especially desert and semi-desert regions, except the highest altitudes and latitudes. They are not as common in the tropics, but genera such as Cephalodromia and Platypygus are known from these regions. Many of these "microbombyliids" have humpbacked thoraces and lack the dense vestiture common in Bombyliidae. Mythicomyiids have until recently not had much attention in the literature. Their small size has caused them to be missed when collecting. Yellow pan trapping and fine-mesh netting in Malaise and aerial sweep nets has resulted in a tremendous amount of undescribed material from many parts of the world. A high amount of diversity of both genera and species exists for this family in Africa, especially northern and southern portions. Approximately 350 species are known (most in the genus Mythicomyia Coquillett). Hundreds more await description.

Because of their extremely small size and curious body shapes, some genera have been at times placed in Acroceridae or Empididae. Originally, taxa were placed in the subfamily Mythicomyiinae in the Empididae. Then later they were transferred to the Bombyliidae, where mythicomyiids have long been treated. Zaitzev (1991) was the first to give characters warranting raising the group to family level. Subsequent workers have followed Zaitzev's lead and treat the group as a separate family. The family is separated from the Bombyliidae by the unbranched wing vein R4+5 (branched in Bombyliidae), the extremely reduced or absent maxillary palpi (present in Bombyliidae), wings held together over the abdomen at rest (held at an angle in Bombyliidae), and the abdominal spiracles being placed in the terga (placed in the pleural membrane in Bombyliidae). Augmenting the morphological characters, it is also a much older lineage than any known Bombyliidae, dating from as far back as the Middle Jurassic (Palaeoplatypygus Kovalev; Callovian: 163–168 mya) with other genera known from the Cretaceous (Procyrtosia Hennig and Proplatypygus Zaitzev). Bombyliidae, on the other hand are not known from any older fossil material than Eocene Baltic amber deposits (Lutetian to Rupelian: 30–52 mya).

Genera

• Acridophagus Evenhuis, 1983
• Ahessea Greathead & Evenhuis, 2001
• Borissovia Evenhuis, 2003
• Carmenelectra Evenhuis, 2003
• Cephalodromia Becker, 1914
• Cyrtisiopsis Séguy, 1930
• Cyrtosia^{[disambiguation needed ]} Perris, 1839
• Doliopteryx Hesse, 1938
• Elachymyia Hall & Evenhuis, 1987
• Empidideicus Becker, 1907
• Eurodoliopteryx Nel, 2006
• Glabellula Bezzi, 1902
• Glella Greathead & Evenhuis, 2001
• Hesychastes Evenhuis, 2002
• Heterhybos Brèthes, 1919
• Leylaiya Efflatoun, 1945
• Mnemomyia Bowden, 1975
• Mythenteles Hall & Evenhuis, 1986
• Mythicomyia Coquillett, 1893
• Nexus Hall & Evenhuis, 1987
• Onchopelma Hesse, 1938
• Palaeoplatypygus Kovalev, 1985
• Paraconsors Hall & Evenhuis, 1987
• Pieza Evenhuis, 2002
• Platypygus Loew, 1844
• Procyrtosia Zaitzev, 1986
• Proplatypygus Hennig, 1969
• Pseudoglabellula Hesse, 1967
• Psiloderoides Hesse, 1967
• Reissa Evenhuis & Baéz, 2001

References

• Evenhuis, N.L., 2002. Catalog of the Mythicomyiidae of the world.Bishop Museum Bulletin in Entomology 10: 1-85. established classification of the family.
• Greathead, D.J. & N.L. Evenhuis, 1997. Family Bombyliidae. In: Contributions to a Manual of Palaearctic Diptera Volume 2 (L. Papp & B. Darvas, eds.): 487-512. Science Herald, Budapest. provide a key to the Palaearctic genera.
• Greathead, D.J. & N.L. Evenhuis, 2001. Annotated keys to the genera of African Bombylioidea (Diptera: Bombyliidae; Mythicomyiidae). African Invertebrates 42: 105-224. good illustrated keys to African genera.
• Zaitzev, V.F. 1991. the phylogeny and system of dipterous insects of the superfamily Bombylioidea (Diptera). Entomologicheskoe Obozreniye 70: 716-736. [English translation, 1992, in Entomological Review 71(4): 94-114.]

External links

• Diptera.info Images
• world catalog of Mythicomyiidae (original and updated catalog as pdf)
• catalog of the Australian species of Mythicomyiidae
• catalog of the fossil species of Mythicomyiidae
• photo of Platypygus ridibundus